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Mixed short/tall tussock grassland
e.g. Chionochloa rigida with Festuca novae-zelandiae
Diagnostic features for identification in the field
Narrow-leaved snow tussock:
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Descriptions of narrow-leaved snow tussock are complicated by the fact that this species is now considered to form a complex with broad-leaved snow tussock (C. flavescens). There are five recognised taxa (including three broad-leaved ones which are described under C. flavescens), which vary in locality and habitat. However, the general description below will differentiate the narrow-leaved members of this complex from the other Chionochloa tussock species.
A large green or tawny green tussock, usually less than 150cm tall. Tussocks may be smaller with narrow leaves, where soil fertility is reduced or at higher altitudes.
Leaves are relatively broad, which droop. They are tough and glossy, and lack a conspicuous midrib above or beneath. The leaves tend to trail on the ground, so that the exposed soil shows a swept effect.
The sheaths are a lighter colour than other species, being pale green to light brown to orange, often purple tinted. Young sheaths are white-hairy, while dead sheaths break crosswise.
The flower heads are borne well above the leaves.
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Fescue tussock:
- An erect fawn tussock 30 - 60cm tall.
- Leaves are rough on the fingers or tongue, rigid, with a sharp point.
- The leaf sheath is fissured to the base.
- The ligule is hairy and asymmetrical.
- Flower stalk is 30 - 60cm long and rough to the touch.
- Flower head is open, with few flowers. The awns are short.
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References:
- Jane, G. 1995. Field guide to grasses. 1. Alpine grasses.
- Mark, A.F. and Adams, N. 1996. New Zealand alpine plants, Methuen.
- Wilson, H.W. 1996. Wild plants of Mount Cook National Park, Manuka Press, Christchurch.
Description of community and habitat
East of the Main Divide, South Island
Narrow-leaved snow tussock grassland is the dominant vegetation cover at mid altitude on mountain slopes south of the Rakaia River, and the hill tops of Banks Peninsula. It's current distribution is a shadow of it's extent prior to European settlement. Tussock size is influenced by the harshness of the climate in which it grows. Where present near sea-level in south-east Otago, tussocks are up to 1.5m tall, but reduce to 0.3m tall at high altitudes.
The abundance and composition of species growing in narrow-leaved snow tussocks is influenced by the degree of disturbance that the grassland has experienced. Mixed short/tall tussock grasslands develop where tall tussock grasslands have been severely opened up by grazing disturbance, resulting in low tall tussock cover, interspersed with short tussocks and an abundance of intertussock herbs and grasses. In Canterbury, mountain cotton daisy (Celmisia spectabilis) is a common associate, especially in more modified grasslands. These open canopied mixed short/tall tussock grasslands are widespread today, especially at mid-altitude, where extensive pastoral development has also taken place.
North Island
Mixed short/tall tussock grasslands (with narrow-leaved snow tussocks) are not present in the North Island.
Other species present
East of the Main Divide, South Island
These open canopied mixed short/tall tussock grasslands often support a diverse array of intertussock herbs and grasses. Mat daisies (Raoulia subsericea) are common, while grasses often include Deyeuxia avenoides, Elymus solandri, and the exotic brown top (Agrostis capillaris) and sweet vernal (Anthoxanthum odoratum). Mouse-eared hawkweed (Hieracium pilosella) is the most widespread hawkweed, although tussock hawkweed (H. lepidulum) and king devil hawkweed (H. praealtum) can also be common. Matagouri (Discaria toumatou) is the most common shrub associated with mixed short/tall tussock grassland, especially where soil fertility has been increased through the aerial application of fertiliser. Native broom (Carmichaelia spp), and Leucopogon colensoi can be locally important. In mid Canterbury, mountain cotton daisy is a common sight in open short/tall tussock grasslands, and is a sign of overgrazing.
Vegetation History
Much of the mountainlands to the east of the Main Divide were covered in forests prior to 700 - 900 years ago. These were largely destroyed by natural and Polynesian fires prior to European settlement, leading to a migration of tussock species into previously forested areas. Initially, short tussock species are thought to have migrated from enclaves within the forest, upslope. This was followed by a slower migration of Chionochloa species (including C. rigida) naturally occupying alpine areas above the timberline, downslope into penalpine, subalpine and even montane environments. Subsequent modification of these tall tussock grasslands in subalpine and montane zones, through pastoral activities (grazing and burning) has lead to a transformation to short tussock grasslands. At mid altitudes, mixed short/tall tussock grasslands commonly occur. They represent a transitionary state between tall tussock grasslands and short tussock grasslands.
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